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8 exocad v SPEAG SEMCAD X Matterhorn v15 troduction. 4 Respiratory Pharmacology Faculty of Medicine, National Heart , Lung Institute Imperial College. 2 Faculty of Veterinary Medicine . In addition, HDAC6 affects antigen- dependent CD4+ T cell activation.Fulwyler developed this in 1965 with his publication in Science. 4 Dolphin Imaging v11. Immunization studies revealed defective cytotoxic activity in vivo in the absence of HDAC6. In this study, we show that HDAC6 contributes to the cytotoxic function of CD8+ T cells. Mack Fulwyler was the inventor of the forerunner to today' s flow cytometers - particularly the cell sorter.